Washington State University

School of the Environment

Lisa Shipley

Associate Director/Professor

Dr. Lisa A. Shipley is a wildlife ecologist in the School of the Environment at Washington State University. Her research interests include foraging ecology and nutrition of mammals, interactions between plants and herbivores at different spatial scales, and ecology and management of wild ungulates. Both free-ranging and captive herbivores, such as mule deer, pronghorn, elk, and pygmy rabbits are used in these studies. Lisa directs the Wild Herbivore Ecology Lab, which includes the Wild Ungulate Facility and the Small Mammal Research Facility at WSU. She is the faculty advisor of the WSU Student Chapter of The Wildlife Society.


  • NATRS 310-Methods of Wildlife Ecology
  • NATRS 446-Wildlife Habitat Ecology
  • NATRS 595-Seminar in Natural Resource Sciences

Research Interests:

  • Foraging ecology of mammals, interactions between plants and herbivores, and ecology and management of wild ungulates
  • The influence of animal size, digestive morphology, and feeding strategy on the diet and habitats selected by herbivores (both free-ranging and captive herbivores, such as woodland
    caribou, blue duikers, porcupines, and pygmy rabbits, are used in these studies).




Selected Publications:

  • Crowell, M.,L. A. Shipley, M. J. Camp,  J. L. Rachlow, J. S. Forbey, and T. R.Johnson. 2016. Selection of food patches by sympatric herbivores in response to concealment and distance from a refuge.  Ecology and Evolution DOI: 10.1002/ece3.1940.
  • Camp, M. J., L. A. Shipley, T. R. Johnson, M. M. Crowell, J. S. Forbey, and J. L. Rachlow.  2015.Modeling tradeoffs between plant fiber and toxins:  A framework for quantifying risks perceived by foraging herbivores. Ecology. 96:3292-3302
  • Stonehouse, K. F., L. A. Shipley, J. Lowe, M. T. Atamian, M. E. Swanson, and M. A. Schroeder. 2015. Habitat selection by sympatric greater sage-grouse and Columbian sharp-tailed grouse.  Journal of Wildlife Management 79:1308-1326.
  • Olsoy, P. J., J. S. Forbey, J. L. Rachlow, J. D. Nobler, N. F. Glenn, and L. A. Shipley.  2015. Fearscapes: mapping functional properties of cover for prey with terrestrial LiDAR.  Bioscience 65:74-80.
  • Rachlow, J. L., R. M. Peter, L. A. Shipley, and T. R. Johnson.  2014.  Sublethal effects of capture and collaring on wildlife: experimental and field evidence. Wildlife Society Bulletin 38:458-465.
  • Ulappa, A. C., R. G. Kelsey, G. G. Frye, J. L. Rachlow, L. A. Shipley, X. Pu, L. Bond, and J. Forbey.  2014. Plant protein and secondary metabolites influence diet selection in a mammalian pecialist herbivore. Journal of Mammalogy 95:834-842.
  • Camp, M. J., J. L. Rachlow, L. A. Shipley, T. R. Johnson, K D. Bockting.  2014. Grazing in sagebrush rangelands in western North American: Implications for habitat quality for a sagebrush specialist,  the pygmy rabbit. The  Rangeland Journal 36:151-159.
  • Elias, B. A., L. A. Shipley, S. McCusker, R. D. Sayler, and T. R. Johnson. 2013. Effects of genetic management  on reproduction, growth, and survival in captive endangered pygmy rabbits (Brachylagus idahoensis).Journal of Mammalogy 94:1282-1292.
  • Ellsworth, E., A. J. Wirsing, L. A. Shipley, and D. L. Murray. 2013.  Do measures of plant intake and digestibility from captive feeding trials align with foraging patterns of free-ranging snowshoe hares? Wildlife Research 40:340-357.
  • Cook, R. C., J. G. Cook, D. J. Vales, B, K. Johnson, S. M. McCorquodale, L. A. Shipley,  R. A. Riggs,  L. Irwin, S. L. Murphie, B. L. Murphie, K. A. Schoenecker, F. Geyer, P. B. Hall, R. D. Spencer, D. A. Immell, D. H. Jackson, B. L. Tiller, P. J. Miller, and L. Schmitz. 2013. Regional and seasonal patterns of nutritional condition and reproduction in elk. Wildlife Monographs 184.
  • Wagoner, S. J., L. A. Shipley, R. C. Cook, and L. Hardesty. 2013   Spring cattle grazing and mule deer nutrition in a bluebunch wheatgrass community.  Journal of Wildlife Management 77:897-907.
  • Camp, M. J., B. A. Woods, J. L. Rachlow, T. R. Johnson, and L. A. Shipley.  2013.  Examining the functional components of cover: The relationship between concealment and visibility in shrub-steppe habitat. Ecosphere 4:1-14.
  • Camp, M. J., J. L. Rachlow, B. A. Woods, T. R. Johnson, and L. A. Shipley.  2012.  When to run and when to hide: The influence of concealment, visibility, and proximity to refugia on perception of risk.  Ethology 118:1010-1017.
  • Shipley, L. A., E. M. Davis, L. A. Felicetti, S. McClean, and J. S. Forbey. 2012. Mechanisms for eliminating monoterpenes of sagebrush by specialist and generalist rabbits.  Journal of Chemical Ecology 38:1178-1189.
  • Tollefson, T. L., L. A. Shipley, W. L. Myers, and N. Dasgupta. 2011. Forage quality’s influence on mule deer fawns.  Journal of Wildlife Management 75:919-928.
  • McCusker, S., L. A. Shipley, T. N. Tollefson, M. Griffin, and E. A. Koutsos. 2011.  Effects of starch and fiber in pelleted diets on nutritional status of mule deer (Odocoileus hemionus) fawns. Journal of  Animal  Physiology and Animal Nutrition 95: 489 – 498.
  • Tollefson, T. L., L. A. Shipley, W. L. Myers, D. H. Keisler, and N. Dasgupta. 2010.  The influence of summer and autumn nutrition on body condition and reproduction in lactating mule deer.  Journal of Wildlife Management 74:974-986.
  • Johnstone-Yellin, T. L., L. A. Shipley, W. L. Myers, and H. S. Robinson.  2009.  To twin or not to  twin? Tradeoffs in litter size and fawn survival in mule deer.  Journal of Mammalogy 90:453-460
  • Shipley, L. A., J. S. Forbey, and B. D. Moore. 2009.  Revisiting the dietary niche: when is a mammalian herbivore a specialist?  Integrative and Comparative Biology 49:274-290.
  • Kendrick, E., L. A. Shipley, A. E. Hagerman, and L. M. Kelly.  2009. Fruit and fibre: The nutritional  value of figs for a small tropical ruminant, the blue duiker (Cephalophus monticola).  Journal of  African Ecology 47:556-566
  • DeGabriel, J. L., B. D. Moore, L. A. Shipley, A. K. Krockenberger, I. R. Wallis, C. N. Johnson, and W. J.  Foley.  2009. Inter-population differences in the tolerance of a marsupial folivore to plant  secondary metabolites.  Oecologia 161:534-548.
  • Shipley, L. A.  2007.  The influence of bite size on foraging at larger spatial and temporal scales by  mammalian herbivores.  Oikos 116:1964-1974.
  • Robbins, C. T., J. K. Fortin, K. D. Rode, S. D. Farley, and L. A. Shipley.  2007.  Optimizing protein intake as a foraging strategy to maximize mass gain in an omnivore.  Oikos 116:1675-1682.
  • Cook, R. C., T. R. Stephenson, W. L. Myers, J. G. Cook, and L. A. Shipley.  2007. Validating predictive models of nutritional condition for mule deer.  Journal of Wildlife Management 71:1934-1943.
  • Thines, N. J., L. A. Shipley, J. H. Bassman, J. K. Fellman, D. S. Mattison, and J. R. Slusser. 2007. Effects of enhanced UV-B radiation on plant chemistry: Nutritional consequences for a specialist and generalist lagomorph.Journal of Chemical Ecology 33:1025-1039.
  • Shipley, L.. A., T. B. Davila, N. J. Thines, and B. A. Elias.  2006. Nutritional requirements and diet choices of the  pygmy rabbit (Brachylagus idahoensis): A sagebrush specialist.  Journal of Chemical Ecology 32:2455-2474.
  • Elias, B. A., L. A. Shipley, R. D. Sayler and R. S. Lamson.  2006. Mating behavior and parental care in captive pygmy rabbits. Journal of Mammalogy  87:921-928
  • Johnstone-Yellin, T. L., L. A. Shipley, and W. L. Myers. 2006. Evaluating the performance of vaginal implant transmitters for capturing neonatal mule deer fawns.  Wildlife Society Bulletin 34:338- 344.
  • Searle, K., T. Vandervelde, N. T. Hobbs, L. A. Shipley, and B. A. Wunder. 2006.  Spatial Context influences patch residence time in foraging hierarchies. Oecologia 148:710-719.
  • Searle, K., N. T. Hobbs, B. A. Wunder, and L. A. Shipley. 2006. . Preferences in a patchy landscapes: the influence of scale-specific intake rates and variance in reward. BehavioralEcology 17:315-323.


School of the Environment
Washington State University
425 Heald Hall
Pullman, WA, 99164-2812

(509) 335-9182


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